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Well, maybe the Twili's powers are what changed the mask. Maybe it changed to suit the Twili's appearance when the first one wore it. About your last question, well I have no clue :).

Well, maybe the ancient tribe weren't Twili to start out with, but were banished because they created and use Majora's mask. Then when they were banished, maybe they changed according to their environment. Also, either the masks powers changed it, the environment changed it, or Onilink is right...