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When I read that, I actually got the impression that the towers are just entrances to a single, much larger facility... similar to how the shrine entrances are pretty small compared to the actual shrines underneath.

I'm not sure if the fifth dungeon will actually be a divine beast... there is a mention of there being some sort of ancient facility beneath Hyrule Castle, so I think it'll be that. If there was a fifth divine beast, then I think it would have been mentioned at some point, but it's possible they...